American video game designer Roger Dearly lives with his pet Dalmatian Pongo in London. One day, Pongo sets his eyes on Perdita, another Dalmatian. After a chase that ends in St James’s Park, Roger discovers that Pongo likes Perdita. Her owner, Anita Campbell-Green, falls in love with Roger when they meet. When they return to Roger’s home, Anita accepts his proposal. They get married along with Perdita and Pongo. Anita works as a fashion designer at the House of de Vil. Her boss, Cruella de Vil, has a deep passion for fur, going so far as to have a taxidermist, Mr. Skinner, skin a white tiger at the London Zoo to make her into a rug for her. Anita, inspired by her Dalmatian, designs a coat made with spotted fur. Cruella is intrigued by the idea of making garments out of actual Dalmatians, and finds it amusing that it would seem as if she was wearing Anita’s dog.

Perdita gets pregnant, and so does Anita. Cruella visits their home and gets excited when she finds out Perdita is expecting. Weeks later, she returns when a litter of 15 puppies are born and offers Roger and Anita £7,500 for them, but they refuse. Enraged, Cruella dismisses Anita and vows revenge against her and Roger. One winter evening, she has her henchmen, Jasper and Horace, break into their home and steal the puppies, while the couple is walking in the park with Pongo and Perdita. Along with 84 other Dalmatians that were previously stolen, they deliver them to her country estate, De Vil Mansion. Cruella asks Skinner to kill and skin them to create her coat.

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